Fruit growth and cell growth-related parameters during fruit development in thinned and un-thinned fruit of ‘Golden Delicious Smoothee’ (GS). (A) Fruit diameter, (B) cell number (layers), (C) relative cell production rate (RCPR) and (D) cell area were measured from bloom to maturity. Closed circles represent un-thinned fruit and open circles represent thinned fruit. Dotted line represents the day of manual thinning (11 DAFB). Error bars represent the standard error of the mean of four biological replicates (n = 4).