Restored root growth in bps1-2 rml1-1 plants reinstates arrest of leaf development. Top row: rml1-1 grown for 28 days on normal growth medium (GM) (left) and or transferred to GSH-supplemented GM at day 10 (right). The small stunted root is rescued by transfer to GSH (+). Bottom row: bps1-2 (left) and bps1-2 rml1-1 (right), both transferred from GM to GM+GHS at 10 days. The bps1 single mutants produced small roots and narrow radialized leaves. The bps1-2 rml1-1 double mutants showed a novel phenotype. The roots enlarged radially, they produced a few enlarged lateral-root-like organs, and arrested leaf primordia accumulated at the shoot apex. Size bars = 1 mm except for bps1-2 rml1-1 roots and leaf primordia, where bars = 0.1 mm.