Table 2 Co-localization of candidate genes revealed using the combined GWAS and QTL analysis

HfDTA

qHfLB1-1

chr1.S_19944812

1

T/C

3.86E-05

Zm00001d028010

Conserved oligomeric Golgi complex subunit 8

HnDTA

qHnDTA1-1

chr1.S_12443338

1

A/C

4.38E-05

Zm00001d027752

SH2 domain protein B

HnDTA

qHnDTA1-1

chr1.S_12443395

1

C/T

5.17E-05

Zm00001d027752

SH2 domain protein B

HnDTA

qHnLB1-1

chr1.S_29513840

1

T/G

8.75E-05

Zm00001d028303

Branched-chain-amino-acid aminotransferase

HnDTA

qHnLB1-1

chr1.S_29517378

1

G/T

3.58E-05

Zm00001d028303

Branched-chain-amino-acid aminotransferase

HnDTA

qHnLB1-1

chr1.S_35770394

1

G/C

1.07E-05

Zm00001d028469

PGG domain-containing protein

HnDTA

qHnDTA1-2

chr1.S_283970301

1

C/T

3.92E-05

Zm00001d034317

Methyltransferase

HnDTA

qHnDTA1-2/qHnDTH1-1

chr1.S_286863933

1

G/A

2.39E-05

Zm00001d034406

25.3 kDa vesicle transport protein

HnDTA

qHnDTA1-2/qHnDTH1-1

chr1.S_286980178

1

T/C

1.07E-05

Zm00001d034409

Hydroxyproline-rich glycoprotein family protein

HnDTA

qHnDTA1-2/qHnDTH1-1

chr1.S_290115635

1

G/A

5.62E-05

Zm00001d034519

Putative acyl-activating enzyme 18 peroxisomal

HnDTH

qHnDTA1-2

chr1.S_283970301

1

C/T

3.15E-05

Zm00001d034317

Methyltransferase

HnDTH

qHnDTA1-2/qHnDTH1-1

chr1.S_286863933

1

G/A

1.29E-05

Zm00001d034406

25.3 kDa vesicle transport protein

HnDTH

qHnDTA1-2/qHnDTH1-1

chr1.S_286980295

1

G/A

6.52E-05

Zm00001d034409

Hydroxyproline-rich glycoprotein family protein

HnLA

qHnDTA1-2

chr1.S_283229666

1

A/G

1.37E-05

Zm00001d034276

Uncharacterized protein

HfDTH

qHnLA2-1

chr2.S_42789389

2

C/T

6.62E-05

Zm00001d003446

Gamma-glutamyltranspeptidase 1

HnDTA

qHnTLN2-1

chr2.S_1084283

2

A/G

8.87E-05

Zm00001d001798

Minichromosome instability12b

HnDTA

qHnTLN2-1

chr2.S_1084664

2

T/C

3.25E-06

Zm00001d001798

Minichromosome instability12b

HnLA

qHnLA2-1

chr2.S_44261048

2

G/A

8.97E-05

Zm00001d003499

E3 ubiquitin-protein ligase ATL6

HnDTA

qHfDTH4-2/qHnDTS4-2

chr4.S_224038847

4

T/G

1.07E-05

Zm00001d053376

AAA-type ATPase family protein / ankyrin repeat family protein

HnDTA

qHfDTH4-2/qHnDTS4-2

chr4.S_224038876

4

G/A

1.07E-05

Zm00001d053376

AAA-type ATPase family protein / ankyrin repeat family protein

HfDTA

qHfDTH5-1

chr5.S_18632762

5

C/G

3.53E-05

Zm00001d013745

Uncharacterized protein

HnLB

qHfDTH5-1

chr5.S_17862164

5

T/A

9.68E-05

Zm00001d000202

Uncharacterized protein

HnLB

qHnDTS6-1

chr6.S_163425227

6

G/C

2.65E-05

Zm00001d038954

2-oxoglutarate and Fe (II)-dependent oxygenase superfamily protein

HfDTA

qHnDTS9-1

chr9.S_48271409

9

A/G

2.47E-05

Zm00001d045962

Uncharacterized protein

HnDTA

qHnDTS9-1

chr9.S_48271409

9

A/G

2.31E-05

Zm00001d045962

Uncharacterized protein

HnDTH

qHnDTS9-1

chr9.S_48271409

9

A/G

3.36E-05

Zm00001d045962

Uncharacterized protein