Fig. 5

Vegetative and reproductive development of the wrinkled aerial organs (wao) tomato mutant. a wao plants cultivated in vitro develop wrinkled leaves and show a moderate curvature of the stem. b-d In greenhouse-grown wao plants, the degree of leaf blade wrinkling depends on the ontogenetic stage. The first two leaves are saddle-shaped (b), the wrinkling decreases in young leaves (c), and increases again in the adult ones, although without being as pronounced as in the first leaves (d). e Adult plant of the wao mutant. f The WT aerial part grafted on its own root (left) grows much more than when grafted on the mutant root (right) indicating that the gene altered in wao also plays a functional role in the root system. g The style of the wao flower is curved and the anthers develop abnormally. h In wao flowers, sepals wrap the internal floral whorls, preventing the expansion of petals in anthesis. i The conversion of ovary to fruit in wao occurs in pre-anthesis. j At the mature green stage the cuticle of wao fruits is wrinkled, giving it a rough texture. k The alteration in cuticle structure most probably explains the tendency to cracking in mature red fruits. Bar (a-f, h, j and k) = 1 cm; Bar (g and i) = 1 mm