Variant leaf morphologies in diploid (2x – left) and tetraploid (4x – right) wild type (A), and transformants (B-E) carrying various constructs. A. Diploid (left) and tetraploid (right) forms of ‘Hawaii 4’. B. Construct E08 2x and 4x transformants displaying elongated petiolules (arrow) and subtle alterations in leaflet shape. Leaflets of tetraploid E08 regenerants were broadened as associated with tetraploidy. C. The leaves of construct F10 transformants have a ruffled appearance, which is magnified in the 4x form. Elongated petiolules are also sometimes present in the 4x form. D. In construct D09 2x transformants, leaves are smaller than wild type, primarily due to narrowing of the leaflets. E. In construct E10 transformants, there is a variable tendency to develop leaves with four or five leaflets (see also Figure 6).