Expression of 5 S rRNA genes linked to 35 S rDNA in plants, their epigenetic modification and regulatory element divergence

Table 2 Relationship between genomic organization and DNA methylation of 5 S genes in the studied species

Species	¹Organization	Number of 5 S loci	¹Position on chromosomes	²mC average	mC range
Artemisia absinthium	L_S	2	(sub-)terminal	46%	34-62%
Tagetes patula	L	7	(sub-)terminal	33%	11-44%
Elachanthemum intricatum	S_L	n.d.	n. d.	39%	31-45%
Helichrysum bracteatum	L	1	(sub-)terminal	34%	16-43%
Helianthus annuus	S	5	pericentromeric	63%	30-96%
Arabidopsis thaliana	S	3	pericentromeric	³79%	³36-95%

¹ The genomic organization and chromosomal position of 5 S rDNA is taken from [12–15, 38] and from the Plant rDNA database [39].
²This work.
³ Methylation levels of coding region were determined by bisulfite sequencing. Data for Arabidopsis genes are from [40]; slightly lower values were reported in [41].

ISSN: 1471-2229