QTLs and candidate gene map. The LHRF_ F3:4segregating population was used. Bins are shown on the right of chromosomes. Distances are in pcM in the LHRF_ F3:4map (LHREF3_1201). QTLs for desiccation (blue), desiccation rate (green), growth (black) and ABA content (pink) are on the left of the chromosomes (Tables 2 and 3). The confidence intervals of the QTLs are indicated by vertical bars. Arrows highlight colocations between desiccation QTLs (blue) and growth QTLs (black) and between desiccation QTLs and ABA QTLs (pink). Gene locations in the QTL confidence interval are indicated close to their corresponding QTLs. Gene codes are detailed in Tables 3 and 4. Genes in bold were mapped by PCR in this study (Additional file 2). The others were mapped by Génoplante http://urgi.versailles.inra.fr/GnpMap/. Genes involved in kernel drying and located in the vicinity of the QTL confidence interval, are indicated in grey. ABA-emb: ABA in embryo (pg/DW); ABA-end: ABA in endosperm (pg/DW); DW: kernel dry matter weight (mg/kernel); FW: kernel fresh matter weight (mg/kernel); %DW = DW/FW× 100; Water: kernel water content (mg/kernel); Rate = (Water/FW × 100)/(thermal time interval); Slope: regression line slope of (Water/FW × 100) as a function of thermal time.